Curriculum Vitae

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Research Interests:

Allometry | Energetics | Muscles as Springs | Clinical Research


Stan L. Lindstedt

Department of Biological Sciences

Northern Arizona University

South Beaver Street

Flagstaff, AZ 86011-5640

Office (928) 523-7524

Lab (928) 523-2439

FAX: (928) 523-7500

Home: 3518 W. Kiltie Loop

Flagstaff, AZ 86001

(928) 779-3605


1970 B.S. (Biology) University of Southern California.

1977 Ph.D.(zoology) University of Arizona. Dissertation director W.A. Calder III.

1977-79 NIH Individual Postdoctoral Fellowship. Dept. of Physiology, University of Arizona, College of Medicine.

1980-81 Swiss NSF Fellowship. Dept. of Anatomy, University of Berne, Switzerland.


1977-79 Research Associate, University of Arizona, College of Medicine

1979-81 International Fellow, University of Berne

1981-1987 Assistant Professor, University of Wyoming

1987-1988 Associate Professor, University of Wyoming

1985-1989 Associate of Zoology (adjunct faculty), Harvard University

1988-1990 Assistant Professor, Northern Arizona University

1990-1994 Associate Professor, Northern Arizona University

1994-1999 Professor of Biology, Northern Arizona University

1999-present Regents’ Professor of Biology, Northern Arizona University

2000-present Professor, Physiological Sciences Program, University of Arizona

academic service:

National: National Science Foundation: Study Section Member (1996-2000); American Heart Association: Member of both the Northwest/Rocky Mountain (1985-88) and Southwest (1992-96) Research Peer Review Committees; Chair (2001-2004) and Treasurer (1993-96) and member of Steering Committee, Comparative Physiology Section of the American Physiological Society; Arizona Disease Control Commission member (1995-1998); Planning Committee, EPA workshop on pharmacokinetics (1988).

University of Wyoming: Chair, General Education (Liberal Studies) Committee, College of Arts and Sciences; Chair, NIH/Biomedical Research Support Grant Committee; Chair, Graduate Selection Committee.

Northern Arizona University: Chair, Biology Department Chair Search Committee; President Search Committee; Provost Search Committee; Interim Co-director (with Richard Posner) of NAU-MBRS program; A&S Dean’s Search Committee; A&S College P&T Committee; University Strategic Planning Council; University Restructuring Committee; Faculty Senate; Ad Hoc academic misconduct inquiry committee; Biology Department Committee on Faculty Status; Biology Department Advisory Committee; Chair, Institutional Animal Use and Care Committee; Academic Excellence Committee; Undergraduate Affairs Committee; Graduate Affairs Committee; Chair, Graduate Admissions Committee, Graduate Advisory Committee; Undergraduate Honors Committee; Library Committee; Chair (1), Department Faculty Search Committees (4); 10 PhD and 12 MS Advisory Committees.

Service as referee:

Journals: American Journal of Physiology (Editorial Board member), Journal of Experimental Biology (Editorial Board member), Nature, Science, Journal of Applied Physiology, Ecology, The American Naturalist, Quarterly Review of Biology, Respiration Physiology, Journal of Theoretical Biology, Oikos, So. African Journal of Zoology, Journal of Mammalogy, National Geographic Science, Comparative Biochemistry and Physiology, Canadian Journal of Zoology, Journal of Comparative Physiology, Physiological Zoology.

Funding agencies: National Science Foundation, National Geographic Society, American Heart Association, Arizona Disease Control Commission, Swiss National Science Foundation, Australian National Science Foundation.

awards and honors:

Invited speaker, Gordon Research Conference on the Biology of Aging, Santa Barbara, 1979

Recognition of outstanding teaching, Department of Zoology and Physiology, University of Wyoming, 1985.

Adjunct appointment as Associate of Zoology, Museum of Comparative Zoology, Harvard University, 1985-89

Invited speaker, National Academy of Sciences, Natl. Res. Council Conference on Pharmacokinetics, Washington, DC, 1986

Keuhn Professorship Research Award, academic year 1986-87

Flinn Foundation Junior Fellow, 1989-91

Keynote Address, Southeast American College of Sports Medicine, Chattanooga, TN 1996.

Faculty Mentor Award. Northern Arizona University, 1998.


1976-77 NSF (DEB 7518576) Dissertation Improvement Grant "Behavioral and physiological coupling of the smallest desert mammal to its physical environment."

1977-79 NIH (AM05738) Individual Postdoctoral Fellowship "Renal function in a small, phyletically primitive mammal." $26,000.

1979-81 Swiss NSF (88765079) International Fellowship "Scaling of pulmonary diffusing capacity in mammals." $36,000.

1984-85 American Heart Assoc. (84-055006) "Erythrocyte transit time through the pulmonary capillaries." $10,000.

1986-88 NSF (EPSCoR) "Limits to aerobic capacity in mammals." $99,500 with state match.

1989-91 American Heart Association, Flinn Foundation Fellowship (FG-30389) "Effects of chronic endurance training on the biochemistry and ultrastructure of the myocardium." $50,000 (direct costs).

1989-93 NIH (RO1 HL41986-01) "Muscle adaptability and respiratory system design." $145,418 (direct costs).

1991-1992 NIH (S15 HL47771) Small Instrumentation Grant $5,000 (direct costs).

1992-1993 NSF (INT 9218487) "Effects of hypoxic training on respiratory muscles' strength and endurance." $6,240 (direct costs).

1993-1996. NSF (IBN 9317527) "Muscle energy balance in vivo: Rattlesnake as a model system." SLL, PI , $64,330 (direct costs).

1993-1997 NSF (DUE 9254280) "Slice of Life: An introductory Biology Course" D. Ebert-May, PI; SLL, Co-PI. $403,643 (total costs).

1993-1997 NIH - MBRS (GM 8215-10) Plasticity of skeletal muscle uptake. SLL, PI $128,000 (direct costs).

1997-2000 NSF (IBN 9714731) "Integrated muscle-system design for high flux at minimal cost" $60,000 (total costs, including REU support).

1998-2002 NIH - MSD (R-25 GM 56931) Plasticity of skeletal muscle oxygen uptake. SLL, sub-section PI $126,000 (direct costs).

2001-2002 NIH (R21 AG18701-01A1) "An exploratory intervention for sarcopenia" SLL, PI ($200,000 direct costs).

Pending - NIH (1RO1 AG022520-01A1) Increasing muscle function in cardio-pulmonary patients $2,964,455 (direct costs), SLL PI


Lindstedt, S.L. and W.A. Calder III (1976). Body size and longevity in birds. Condor 78:91-94.

Altschuler, E.M., R.B. Nagle, E.J. Braun, S.L. Lindstedt and P.H. Krutzsch (1979). Morphological study of the desert heteromyid kidney with emphasis on the genus Perognathus. Anat. Record 194:461-468.

Lindstedt, S.L. (1980). Energetics and water economy of the smallest desert mammal. Physiol. Zool. 53:82-97.

Hoppeler, H., O. Mathieu and S.L. Lindstedt (1980). Scaling structural parameters of oxygen consumption against VO2max and body mass. In, Exercise Bioenergetics and Gas Exchange P. Cerretelli & B.J. Whipp (eds.), Elsevier/North-Holland, Amsterdam, pp. 129-135.

Lindstedt, S.L. (1980). The smallest insectivores: Coping with scarcities of energy and water. In: Comparative Physiology, K. Schmidt-Nielsen, L. Bolis & C.R. Taylor (ed.), Cambridge Univ. Press.

Lindstedt, S.L. (1980). Regulated hypothermia in the desert shrew. J. Comp. Physiol. 137:173-176.

Hoppeler, H., O. Mathieu, E.R. Weibel, R. Krauer, S.L. Lindstedt and C.R. Taylor (1981). Design of the mammalian respiratory system. VIII. Capillaries in skeletal muscles. Resp. Physiol. 44:129-150.

Lindstedt, S.L. and Calder W.A. III (1981). Body size, physiological time, and longevity of homeothermic animals. Quart. Rev. Biol. 56:1-15.

Mathieu, O., R. Krauer, H. Hoppeler, P. Gehr, S.L. Lindstedt, R.McN. Alexander, C.R. Taylor and E.R. Weibel (1981). Design of the mammalian respiratory system. VII. Scaling mitochondrial volume in skeletal muscle to body mass. Resp. Physiol. 44:113-128.

Hoppeler, H., S.L. Lindstedt, H. Claasen, C.R. Taylor, O. Mathieu and E.R. Weibel (1984). Scaling mitochondrial volume in heart to body mass. Resp. Phys. 55:131-137.

Lindstedt, S.L. (1984). Pulmonary transit time and diffusing capacity in mammals. Am. J. Physiol. 246:R384-R388.

Hoppeler, H., S.L. Lindstedt, E. Uhlmann, A. Niesel, L.M. Cruz-Orive and E.R. Weibel (1984). Oxygen consumption and the composition of skeletal muscle tissue after training and inactivation in the European woodmouse (Apodemus sylvaticus). J. Comp. Physiol. B. 155:51-61.

Hoppeler, H., H. Howald, K.E. Conley, S.L. Lindstedt, H. Claasen, P. Vock and E.R. Weibel (1985). Endurance training in humans: Aerobic capacity and structure of skeletal muscle. J. Appl. Physiol. 59:320-327.

Hoppeler, H. and S.L. Lindstedt (1985). Malleability of skeletal muscle tissue in overcoming limitations: Structural elements. J. Exp. Biol. 115:355-364.

Lindstedt, S.L. (1985). Birds. In: F.A. Lints (ed.) Non-mammalian Models for Research in Aging. S. Karger, Basel. p. 1-21.

Lindstedt, S.L. and M.S. Boyce (1985). Seasonality, body size and survival time in mammals. Amer. Natur. 125:873-878.

Lindstedt, S.L., H. Hoppeler, K.M. Bard and H.A. Thronson-Jr. (1985). Estimate of muscle shortening rate during locomotion. Am. J. Physiol. 249:R699-R703.

Graves, B.M., D. Duvall, M.B. King, S.L. Lindstedt and W.A. Gern (1986). Initial den location by neonatal prairie rattlesnakes: Functions, causes and natural history in chemical ecology. Chemical Signals in Vertebrates. 4:285-304.

Lindstedt, S.L., B.J. Miller and S.W. Buskirk (1986). Body size, time, and home range size in mammals. Ecology 67:413-418.

Thronson H.A. Jr. and S.L. Lindstedt (1986). The graying of American Astronomy. J. Astron. Soc. Pacific 98:941-947.

Hoppeler, H., J.H. Jones, S.L. Lindstedt, K.E. Claassen; K.E. Longworth, C.R. Taylor, R. Straub and A. Lindholm (1987). Relating maximal oxygen consumption to skeletal muscle mitochondria in horses. p. 278-290 In: J. P. Gillespy, D.H. Snow and R.J. Rose (eds.), Equine Physiology II. Granta Publ., Cambridge.

Lindstedt, S.L. (1987). Allometry: Body size constraints of animal design. Pages 65-79 in: Pharmacokinetics in Risk Assessment. National Academy Press, Washington, DC.

Karas, R.H., C.R. Taylor, J.H. Jones, S.L. Lindstedt, B.R. Reeves and E.R. Weibel (1987). Adaptive variation in the mammalian respiratory system: VII. Flow of oxygen across the pulmonary gas exchanger. Respir. Physiol. 69:101-115.

Lindstedt, S.L. and J.H. Jones (1987). Symmorphosis: the concept of optimal design. In, M. Feder, A.F. Bennett, W. Burrgren and R. Huey (ed.), New Directions in Ecological Physiology. Pages 289-309, Cambridge University Press.

Lindstedt, S.L. and S.D. Swain (1988). Body size as a constraint of design and function. Pages 93-105 in Evolution of Life Histories of Mammals. M. S. Boyce (ed.), Yale Univ. Press.

Lindstedt, S.L. and D.J. Wells (1988). Skeletal muscle mitochondria: The aerobic gate? Pages 207-213 in Oxygen Transfer From Atmosphere To Tissues, Advances in Experimental Medicine and Biology, Vol. 227. N. C. Gonzalez and M. R. Fedde (ed.), Plenum Press, New York.

Lindstedt, S.L., D.J. Wells, J.H. Jones, H. Hoppeler and H.A. Thronson Jr. (1988). Limitations to Aerobic Performance in Mammals: Interaction of Structure and Demand. Int. J. Sports Med. 9:210-217.

Buskirk, S.W. and S.L. Lindstedt (1989). Sex biases in trapped samples of Mustelidea. J. Mammal. 70:88-97.

Kayar, S.R., H. Hoppeler, S.L. Lindstedt, H. Claassen, J.H. Jones, B. Essen-Gustavsson and C.R. Taylor (1989). Total mitochondrial volume in relation to aerobic capacity of horses and steers. Pfluegers Arch. Eur. J. Physiol. 41:343-347.

Lindstedt, S.L., J.F. Hokanson, D.J. Wells, S.D. Swain, H. Hoppeler and V. Navarro (1991). Running energetics in the pronghorn antelope. Nature (cover article) 353:748-750.

Kayar, S.R., H. Hoppeler, R.B. Armstrong, M.H. Laughlin, S.L. Lindstedt, J.H. Jones, K.E. Conley and C.R. Taylor (1992). Estimating transit time for capillary blood in selected muscles of exercising animals. Pfluegers Arch. 421:578-584.

Lindstedt, S.L. (1992). Oxygen supply to the tissues: Keeping up with demand. In E. Bicudo (ed.) Oxygen Transport to the Tissues: Adaptations to Extreme Environments. CRC Press, Boca Raton, FL.

Jones, J.H. and S.L. Lindstedt (1993). Limits to maximal performance. Ann. Rev. Physiol. 55:547-569.

Lindstedt, S.L. and R.G. Thomas (1994). Exercise performance in mammals: An allometric perspective. In J.H. Jones (ed.) Comparative Vertebrate Exercise Performance, Advances in Veterinary Science and Comparative Medicine 38b: 191-217, Academic Press, NY.

Lindstedt, S.L., R.G. Thomas and D.E. Leith (1994). Does peak inspiratory flow contribute to setting ? A test of symmorphosis. Respir. Physiol. 95:109-118

Lindstedt, S.L. and H. Hoppeler (1995). Fatigue and the design of the respiratory system. In D.G. Stuart (ed.) Neural and Neuromuscular aspects of muscle fatigue. Plenum Press, New York.

Schaeffer, P.J., K.E. Conley and S. L. Lindstedt (1996). Structural correlates of speed and endurance in skeletal muscle: The rattlesnake tailshaker muscle. J. Exp. Biol. 198: 351-358.

Rome, L.C., D.A. Syme, S. Hollingsworth, S.L. Lindstedt and S.M. Baylor (1996) The whistle and the rattle: The design of sound producing muscles. Proc. Nat. Acad. Sci. 93: 8095-8100.

K.E. Conley and S.L. Lindstedt. (1996) Rattlesnake tail-shaking: minimal cost per twitch in striated muscle. Nature 383:71-73.

Rome, L.C. and S.L. Lindstedt (1997). Mechanical and metabolic adaptation of the muscular system. In, W.H. Dantzler, ed. Handbook of Physiology, Section 1: Vol. II Comparative Physiology, pages 1587-1652. American Physiological Society, Bethesda, MD.

Conley, K.E. and S.L. Lindstedt (1997). Balancing energy supply and demand for sound production and flight. In, E.R. Weibel, C.R. Taylor and L. Bolis, ed. Optimization in Biological Design: Controversies About Symmorphosis. Cambridge Univ. Press.

Brown, R.P., M.D. Delp, S.L. Lindstedt, L.R.Rhomberg and R.P. Beliles. (1997). Physiological parameter values for physiologically based pharmacokinetic models. Toxicology and Industrial Health 13: 407-484

Lindstedt, S. L. (1997) Design of cells for metabolism, Overview. In, E.R. Weibel, C.R. Taylor and L. Bolis, ed. Optimization in Biological Design: Controversies About Symmorphosis. Cambridge Univ. Press.

Leuenberger, M., A. Grünenfelder, H. Hoppeler, R. Favier, H. Speilvogel, S.L. Lindstedt, G. Ferretti and B. Kayser. (1997). Funktionelle und strukturelle Anpassungen an ein Dauerleistungstraining ober- oder unterhalb der 4 mM/l Laktatschwelle. Schweizerische Z. für Sportmedizin und Sporttraumatologie 45: 97-101.

LaStayo, P.C. and S.L. Lindstedt (1997). Exercise training at high altitude: How do the respiratory muscles respond? Acta Andina 6:29-35.

Hoppeler H, B Kayser, SL Lindstedt, C Boesch, M Kushmerick, A Draeger and F Booth (1997) Recommendations for muscle research in space. Int J Sports Med 1997 Suppl 4:S280-S282

Lindstedt, S. L., T. McGlothin, E. Percy and J. Pifer. (1998) Task-specific design of skeletal muscle: Balancing muscle structural composition. Comparative Biochemistry and Physiology Part B 120:35-40.

Rome, L.C. and S.L. Lindstedt (1998). The Quest for Speed: Muscles built for high frequency contractions. News in the Physiological Sciences 13:261-268 (cover article).

R.G. Thomas, P.C. LaStayo, H. Hoppeler, R. Favier, G. Ferretti, B. Kayser, D. Desplanches, H. Spielvogel and S. L. Lindstedt. (1998) Exercise training in chronic hypoxia has no effect on ventilatory muscle function in humans. Respiration Physiol. 112:195-202.

LaStayo, P.C., T.E. Reich, M. Urquhart, H. Hoppeler and S.L. Lindstedt. (1999) Can improvements in muscle strength be uncoupled from demand for energy during training? Am. J. Physiol. 45:R611-R615.

LaStayo, P.C., D.J. Pierotti, J. Pifer, H. Hoppeler and S.L. Lindstedt. (2000) Eccentric ergometry: Increases in locomotor muscle size and strength at low exercise intensities. Am. J. Physiol. 278:R1282-1288.

Reich, T.E., S.L. Lindstedt, P.C. LaStayo, and D. J. Pierotti. (2000) Is the spring quality of muscle plastic? Am. J. Physiol. 278: R1661-1666.

Kemper W.F., Lindstedt S.L., Hartzler L.K., Hicks J.W and K.E. Conley. (2001) Shaking up glycolysis: Sustained, high lactate flux during aerobic rattling. Proc. Nat. Acad. Sci. 98:723-728.

P. J. Schaeffer, D.J. Wells J.F. Hokanson and S.L. Lindstedt. (2001) Cold exposure increases running and cost of transport in goats. Am. J. Physiol. 280: R42–R47, 2001.

Lindstedt, S.L. and K.E. Conley. Human aerobic performance: Too much ado about limits to . (2001) Journal of Experimental Biology 204: 3195-3199.

Hoppeler, H, R.G. Boutilier and S.L. Lindstedt, editors (2001) Physiological Limits to Human Performance, J. Exp. Biol. Vol 204.

Lindstedt, S.L., P.C. LaStayo and T.E. Reich. When active muscles lengthen: Properties and consequences of eccentric contractions. (2001) News in the Physiological Sciences 16: 256-261.

Lindstedt, S.L. and P.J. Schaeffer. Allometry: How does body size impact design and function of vertebrates? (2002) Laboratory Animals: 36:1-19.

Lindstedt, S.L., T. E. Reich, P. Keim and P. C. LaStayo (2002) Do muscles function as adaptable locomotor springs? J. Exp. Biol. 205:211-2216.

Conley, K.E. and S.L. Lindstedt. (2002) Energy saving mechanisms in muscle: The minimization strategy. J. Expl. Biol. 205: 2175-2181.

Moon, B.R., K.E. Conley, S.L. Lindstedt and M.R. Urquhart (2003) Minimal shortening in a high-frequency muscle. J. Expl. Biol. 206: 1291-1297.

LaStayo, P.C., G. Ewy, D. Pierotti, R. Johns, S.L. Lindstedt (2003) The Positive Effects of negative work: Increased muscle strength and decreased fall-risk in a frail-elderly population. J. Gerontology 58A: 419-424.

LaStayo, P.C., J. Woolf, M.D. Lewek, L. Snyder-Mackler, T. Reich, S.L. Lindstedt.  (2003) Eccentric muscle contractions: Their contribution to injury prevention rehabilitation and sport.  J Ortho Sport Phys Ther, 33: 557-571.

Schaeffer, P.C., J.J. Villarin and S.L. Lindstedt (2003) Chronic cold exposure increases skeletal muscle oxidative structure and function in Monodelphis domestica, a marsupial lacking brown adipose tissue. Physiological and Biochemical Zoology 76: 877-887.

Villarin, J.J., P.J. Schaeffer, R.A. Markle, S.L. Lindstedt (2003) Potential contribution of liver tissue to aerobic evergy balance following chronic cold exposure in the marsupial, Monodelphis domestica. Comp. Biochem. Physiol. A 136: 621-630.


Lindstedt, S.L. and J.H. Jones (1980). Desert Shrewd. Natural History 89:45-53.

Lindstedt, S.L. and H. Hoppeler (1984). A marvelous machine: What you need to know about muscle. Bicycling 24:28-31.

Hoppeler, H. and S.L. Lindstedt (1984). A revolution in endurance training. Bicycling 25:40-46.

Lindstedt, S.L. (1993, editorial invitation). Endurance Physiology. Yearbook of Science and Technology. McGraw-Hill, NY, 153-155.

INVITED LECTURES (excluding home institutions):

1977: Zoology Department Seminars (2) Duke University, Durham, NC

1978: International Conference on Comparative Physiology, Crans, Switzerland.

1979: Gordon Research Conference, The Biology of Aging, Santa Barbara, CA.

        Physiology and Biophysics Department Seminar, University of Illinois, Champaign, IL.

1980: Physiology Department Seminar, Michigan State University, East Lansing, MI.

                            Biology Department Seminar, University of California, Riverside, CA.

1983: Zoology Department Seminar, Colorado State University, Ft. Collins, CO.

1984: Biology Department Seminar, University of California, Los Angeles, CA.

        Physiology and Biophysics Department Seminar, University of Louisville, KY.

                            Physiology Department Seminar, School of Medicine and Dentistry, SUNY, Buffalo.

1985: Museum of Comparative Zoology and Concord Field Station Department Seminars (2) Harvard University, Cambridge.

        Institut de Zoologie et d'ecologie animale Department Seminar, University of Lausanne, Switzerland.

        Zoology Department Seminar, University of Connecticut, Storrs, CT.

        Cell and Developmental Biology Department Seminars (2), University of California, Irvine, CA.

        Physiology Department Seminar, College of Medicine, University of Arizona, Tucson, AZ.

1986: National Research Council, National Academy of Sciences, Washington, DC.

        Special Conference: New directions in physiological ecology, National Science Foundation, Washington, DC

        Workshop on allometry and pharmacokinetics, Environmental Protection Agency, Duluth, MN.

        Fall meeting, American Physiological Society, Symposium on Comparative Physiology.

1987: Evolutionary, Population and Organismal Biology Department Seminar, University of Colorado, Boulder, CO.

        Biology Department Seminar, University of California, Riverside, CA.

1988: Physiology Program Seminar, University of California, Davis, CA.

        Task force on pharmacokinetics and risk assessment. Environmental Protection Agency, Ashville, NC.

        Anatomy Department Seminar, College of Medicine, University of Berne, Switzerland.

1989: Ecology and Evolutionary Biology Department Seminar, University of Arizona, Tucson, AZ.

        International Union of Physiological Sciences, Symposium Lecture, Helsinki, Finland.

        Engineering Foundation Conference on Biomechanics and Neural Control of Movement, Potosi, MO.

                            Biology Department Seminar, Idaho State University, Pocatello, ID.

                            Biology Department Seminar, Boise State University, Boise, ID.

1990: Department of Physiology, College of Medicine, University of Arizona, Tucson, AZ.

1991: American Association of Anatomists, Annual Meeting, Chicago, IL.

        International Congress of Comparative Physiology and Biochemistry, Tokyo.

        Department of Anatomy Seminar, University of Chicago, IL.

                            International Symposium, Oxygen Transport to the Tissues, Sao Paulo, Brazil.

1992: Workshop on Physiologically Based models for Pharmacokinetics, Colorado State University, Ft. Collins, CO.

                            South American Conference on High Altitude Physiology, La Paz, Bolivia.

                             American Physiological Society, Fall Meeting, (two papers) Colorado Springs, CO.

1993: Zoology Department Seminar, University of Washington, Seattle, WA

        10th Annual meeting of the Australian Comparative Physiologists. Hobart, Tasmania.

        Graduate Course in Comparative Physiology (2 lectures) University of California, Irvine.

1994: Department Seminar, Physiology Department, University of Arizona College of Medicine.

        Retirement Symposium in honor of E.R. Weibel. University of Bern, Switzerland.

        Graduate course lecture, comparative physiology, University of Washington, Seattle.

1994: Department Seminar, Biology Department, University of Wollongong, Australia.

        Neural and neuromuscular aspects of muscle fatigue. Invited lecture, Miami, FL.

1995: Life Sciences Seminar, Arizona State University, West. Phoenix, AZ.

        International symposium on Optimality in Biology. 12th International Conference on Comparative Physiology. Ascona, Switzerland.

        AALAS National Meeting symposium speaker. Baltimore, MD.

1996: Keynote Address, Southeast American College of Sports Medicine, Chattanooga, TN.

        Stanford University, Hopkins Marine Station Seminar. Stanford, CA.

        Second World Congress of High Altitude Medicine and Physiology. Cuzco, Peru.

        ESA (European Space Agency)-NASA, Workshop on muscle research in space. Bern, Switzerland.

        Special symposium in tribute to Professor C.R.Taylor. Harvard University, Cambridge.

        Fall meeting of the American Physiological Society. Vancouver, B.C.

1997: XXXIII International Congress of Physiological Sciences, St. Petersburg, Russia.

        11th Annual Roland D. Pinkham Basic Science Lectureship, Seattle, WA.

1998: Arizona Chapter - Society for Neurosciences, Phoenix

                            First Annual Symposium of Sport Science, High Altitude Training Center, Flagstaff

1999: Physiology Department Seminar, University of Arizona College of Medicine.

2000: Limits to Human Performance, special theme conference of the Journal of Experimental Biology, Flagstaff, AZ

        Sarver Heart Center, Arizona Health Sciences Center, Tucson, AZ.

2001: International Union of Physiological Sciences, Satellite symposium, Wollongong, NSW, Australia.

2002: Department Seminar, California Polytechnic University, Pomona, CA

        "Frontiers in Physiology" special seminar series, University of California, Davis, CA

        IGERT seminar series, Arizona State University, Tempe, AZ

        Department Seminar, Kinesiology, University of Illinois, Chicago

2003: Respiratory Sciences Department Seminar, Arizona Health Sciences Center, Tucson, AZ

        Physiology group seminar, Mayo Clinic Rochester, MN

        Bioengineering Seminar, University of Arizona, Tucson, AZ

2004: Department seminar, University of California, Riverside.


Courses taught at NAU

UC 101 (Freshman Colloquium)

Bio 100 and Bio 180 (Non-majors Freshman Biology)

Bio 182 (Unity of Life: Organismal Biology)

Bio 190 (Freshman Animal Biology)

Bio 425 (Animal Physiology)

Bio 540 (Comparative Physiology)

Bio 673 (Physiological Ecology)

Bio 577 (Concepts in Ecology)

Bio 599 (Animal Locomotion)

Bio 498 and 698 (Seminars, many sections, including: Bioenergetics, Allometry, Muscle plasticity, Size and shape in Biology)